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Flyfishing Entomology
 

 

          Entomology Bookcase
 


The last several years have seen increasing interest in entomology, particularly from those who have recently started flyfishing.  At least, that seems to be the case, based on postings from various flyfishing forums.  However, there has not been much comparative analysis of the various flyfishing entomology books.  Hopefully, this article will provide the reader with sufficient information to make more-informed choices.

First, let’s talk about categories of information: 

  • Some books cover most aquatic insect orders, some even cover other aquatic vertebrates, while others focus on a single aquatic insect order, mayflies for example.
  • Some books cover only the immature (nymphal/larval/pupal) life stages, whereas others also cover the adult life stage (or stages).
  • The (taxonomic) level (of description) for books covering multiple insect orders tends to not be below the family (or genus) level.  Conversely, the level of description (for books covering a single order or insects) is more likely to go down to species level.
  • Most books are rich in illustration.  This may include drawings or photos, and in black & white and/or color. 
  • Some books deal with imitation by suggesting patterns, presentation techniques, and emergence timing, while others don’t.
  • Also, these books can vary greatly in such other characteristics as: their scope of geographic coverage; the degree to which their taxonomy has become out-of-date (largely a function of publication date); their availability new (still being published); and their size as measured in number of pages.

So, in order to present all this information to the reader in a consistent manner, use of a  matrix was chosen.  The column headings include the (above) information categories, and the rows are populated with descriptive information from my the entomology books on my  bookshelf.  The books are listed alphabetically in title order.

Also, Topic Organization (below the matrix column for each individual book) should give you a pretty good feel for the nature of the book's content.  Clicking on the "blue" title of a book listed in the Information Summary will take you directly to the detailed information for that book.   

Information Summary

 

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American Stoneflies     •     • •     •     •     • NA 126 • 1998
Angler's Guide To Aquatic Inv. • • • •   • •     • • • •     • NA 182 • 1995

An Intro to the Aquatic Insects

 • • • •   • •   •   •           NA 862 • 1996
Aquatic Entomology • • • • • • • •     •           NA 448 • 1981
Caddisflies   •       • •     • • • • • • • NA 336 • 1981
Guide to Aquatic Inv. of the UM • • • • • •   •     •           UM 207 • 2004
Hatch Guide For Western • • • •   • •   •   •   • • • • W 196 • 1997
Hatch Guide For New England • • • •   • •     •     • • • • NE 271 • 2000
Hatches II •         • •     • • • • • • • NA 335 • 1986
Mayflies (Fauceglia) •         • •     •     • •   • EM 196 • 2005
Mayflies of Michigan T. Stream •         • •     • • • •     • MI 139 • 1962
Mayflies the Angler and the Tr. •         • •     • • • • • • • NA 188   1989
Mayflies (Knopp/Cormier) •         • •     • • • • • • • NA 336 • 1997
Nymphs • • • •   •       • •     • • • NA 339   1973
Selective Trout • • •     • •     • • • • •   • NA 184 • 1971
The Caddisfly Handbook   •       • •     • •   • •   • NA 133 • 1998
The Comp. Book Of W. Hatches • • • •   • •   •   • • • • • • W 223 • 1981
The Hatches Made Simple • • •     • •     • • • • • • • NA 261 • 2002
Western Mayfly Hatches •         • •   •   •   • • • • W 267 • 2004


Geographic coverage legend:
   EM (East & Midwest), MI (Michigan), NA (North America),
   NE (New England), UM (Upper Midwest), W (West)

Individual Books

 

American Stoneflies: A Photographic Guide to the Plecoptera by Bill Stark, Stan Szczytko, and Riley Nelson

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    •     • •     •     •     • NA 126 • 1998
Topic organization:

  Introduction
    What Are Stoneflies?
    Stonefly Anatomy
    Stonefly Biology
    Collecting Stoneflies
    Stonefly Classification and Identification
    Pictorial Key to Stonefly Families
  Stoneflies and Trout
  Family Pteronarcyidae
  Family Peltoperlidae
  Family Perlidae
  Family Perlodidae
  Family Chloroperlidae
  Family Capniidae
  Family Leuctridae
  Family Taeniopterygidae
  Family Nemouridae

An Angler’s Guide to Aquatic Insects And Their Imitations by Rick Hafele and Scott Roederer

 Order Stage Level Illustration Imitation      Other

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• • • •   • •     • • • •     • NA 182 • 1995
Topic organization:

  "Bugs" Is Not a Four-Letter Word
  Fly Patterns: Theory and Design
  Fishing Strategies
  Beginning Identification
  Mayflies: Order Ephemeroptera
  Stoneflies: Order Plecoptera
  Dragonflies and Damselflies: Order Odonata
  Waterbugs: Order Hemiptera
  Alderflies and Dobsonflies: Order Megaloptera
  Caddisflies: Order Trichoptera
  True Flies: Order Diptera
  Beetles: Order Coleoptera
  Aquatic Moths: Order Lepidoptera

An Introduction to the Aquatic Insects Of North America, edited by R. W. Merritt and K. W. Cummings

 Order Stage Level Illustration Imitation      Other

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(update scheduled for early 2007)

 • • • •   • •   •   •           NA 862 • 1996
Topic organization:

  Introduction
  General Morphology of Aquatic Insects
  Design of Aquatic Insect Studies
  Aquatic Insect Respiration
  Habitat, Life History and Behavioral Adaptations
   of Aquatic Insects
  Ecology and Distribution of Aquatic Insects
  Use of Aquatic Insects in Biomonitoring
  Phylogenetic Relationships and Evolutionary Adaptations
   of Aquatic Insects
  General Classification and Key to the Orders
   of Aquatic and Semiaquatic Insects
  Aquatic Collembola
  Ephemeroptera
  Odonata
  Semiaquatic Orthoptera
  Plecoptera
  Aquatic and Semiaquatic Hemiptera
  Megaloptera and Aquatic Neuroptera
  Trichoptera Families
  Trichoptera Genera
  Aquatic and Semiaquatic Lepidoptera
  Aquatic Coleoptera
  Aquatic Hymenoptera
  Aquatic Diptera
    Larvae of Aquatic Diptera
    Pupae and Adults of Aquatic Diptera
  Tipulidae
  Culicidae
  Simuliidae
  Chironomidae

Aquatic Entomology: The Fishermens Guide and Ecologists Illustrated Guide to Insects and Their Relatives by W. Patrick McCafferty

 Order Stage Level Illustration Imitation      Other

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• • • • • • • •     •           NA 448 • 1981
Topic organization:

  BACKGROUND
    Introduction and Organization
    Identifying Aquatic Insects
    Living in Water
    From Field to Laboratory
    Arthropods and Other Common Freshwater
     Macroinvertebrates
  AQUATIC INSECTS
    Key to the Orders and Stages of Aquatic
     Insects
    Mayflies (ORDER EPHEMEROPTERA}
    Damselflies and Dragonflies (ORDER ODONATA)
    Stoneflies (ORDER PLECOPTERA)
    Water bugs (ORDER HEMIPTERA)
    Fishflies, Dobsonflies, and Alderflies
     (ORDER MEGALOPTERA)
    Spongillaflies (ORDER NEUROPTERA, FAMILY SISYRIDAE)
    Water Beetles (ORDER COLEOPTERA)
    Caddisflies (ORDER TRICHOPTERA)
    Aquatic Caterpillars (ORDER LEPIDOPTERA)
    Midges, Mosquitoes, Aquatic Gnats and Flies
     (ORDER DIPTERA)
  OTHER INSECTS ASSOCIATED WITH AQUATIC
     ENVIRONMENTS
    Shore-Dwelling Insects
    Insects Associated With Emergent Vegetation
    Tree Hole and Plant Cup Residents
    Diving Wasps (ORDER HYMENOPTERA)
  OTHER FRESHWATER ARTHROPODS
    Freshwater Springtails (ORDER COLLEMBOLA)
    Common Freshwater Crustaceans
    Common Freshwater Arachnids

Caddisflies by Gary LaFontaine

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  •       • •     • • • • • • • NA 336 • 1981
Topic organization:

  TYING AND FISHING CADDISFLY IMITATIONS
    Anticipating a Caddisfly Hatch
    Creating the Effective Pattern
    Other Effective Caddisfly Imitations
    Fishing the Larval Imitations
    Fishing the Pupal Imitations
    Fishing the Adult Imitations
    Stillwater Caddisflies
  THE BIOLOGY OF CADDISFLIES
    The Biology of Caddisflies
    Caddisfly Habitat
      (The Distribution of Common Caddisfly Genera
       of North American Trout Streams)
    Caddisflies as a Food Resource for Trout
    Insect Listing
      (Emergence and Egg-Laying Charts)
    Collecting and Identifying Caddisflies
      (Checklist of North American Caddisfly Genera)

Guide to Aquatic Invertebrates of the Upper Midwest by R.W. Bouchard

 Order Stage Level Illustration Imitation      Other

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(available for free download)

 • • • • • •   •     •           UM 207 • 2004
Topic organization:

  INTRODUCTION
  AQUATIC INVERTEBRATES
    Key to Aquatic Invertebrates
    Non-Insect Invertebrate Descriptions
    Other Commonly Encountered Invertebrates Not
     Included in the Key
  INSECTA (Aquatic Insects)
    Key to Insecta Orders
  EPHEMEROPTERA (Mayflies)
    Key to Ephemeroptera Families (Larvae)
    Ephemeroptera Family Descriptions
  ODONATA (Dragonflies & Damselflies)
    Key to Odonata Families (Larvae)
    Odonata Family Descriptions
  PLECOPTERA (Stoneflies)
    Key to Plecoptera Families (Larvae)
    Plecoptera Family Descriptions
  HEMIPTERA (Aquatic and Semiaquatic True Bugs)
    Key to Hemiptera Families (Adults)
    Hemiptera Family Descriptions
  MEGALOPTERA (Fishflies, Alderflies & Dobsonflies)
    Key to Megaloptera Families (Larvae)
    Megaloptera Family Descriptions
  NEUROPTERA (Spongillaflies)
    Neuroptera Family Descriptions
  TRICHOPTERA (Caddisflies)
    Key to Trichoptera Families (Larvae)
    Trichoptera Family Descriptions
  LEPIDOPTERA (Aquatic Moths)
    Lepidoptera Family Descriptions
  COLEOPTERA (Aquatic Beetles)
    Key to Coleoptera Families (Larvae)
    Key to Coleoptera Families (Adults)
    Coleoptera Family Descriptions
  Diptera (Aquatic & Semiaquatic True Flies)
    Key to Diptera Families (Larvae)
    Diptera Family Descriptions

Hatch Guide For Western Streams by Jim Schollmeyer

 Order Stage Level Illustration Imitation      Other

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• • • •   • •   •   •   • • • • W 196 • 1997
Topic organization:

  Introduction
  Water Types and Fishing Techniques
  Mayflies
    Blue-Winged Olive
      (Family: Baetidae, Genus: Baetis) Swimmer
    Speckle-Wing Quill
      (Family: Baetidae, Genus: Callibaetis) Swimmer
    Ameletus
      (Family: Ameletidae, Genus: Ameletus) Swimmer
    Black or Gray Drake
      (Family: Siphlonuridae, Genus: Siphlonurus) Swimmer
    Pale Morning Dun
      (Family: Ephemerellidae, Genus: Ephemerella) Crawler
    Western Green Drake
      (Family: Ephemerellidae, Genus: Drunella) Crawler
    Flav
      (Family: Ephemerellidae: Genus: Drunella) Crawler
    Slate-Winged Olive
      (Family: Ephemerellidae, Genus Drunella) Crawler
    Mahogany Dun
      (Family: Leptophlebiidae, Genus: Paraleptophlebia)
      Crawler
    Trico
      (Family: Leptohyphidae, Genus: Tricorythodes) Crawler
    Western March Brown
      (Family: Heptageniidae, Genus: Rhithrogena) Clinger
    Pale Evening Dun
      (Family: Heptageniidae, Genus: Heptagenia) Clinger
    Yellow Quill
      (Family: Heptageniidae, Genus: Epeorus) Clinger
    Blue-Winged Red Quill
      (Family: Heptageniidae, Genus: Cinygmula) Clinger
    Brown Drake
      (Family: Ephemeridae, Genus: Ephemera) Burrower
    Big Yellow May
      (Family: Ephemeridae, Genus: Hexagenia) Crawler
  Caddisflies
    Green Rock Worm
      (Family: Rhyacophilidae, Genus: Rhyacophila)
      Free-Living Caddis
    Spotted Sedge
      (Family: Hydropsychidae, Genus: Hydropsyche) Caddis
      Net-Spinning Caddis
    Glossosoma
      (Family: Glossosomatidae, Genus: Glossosoma) Caddis
      Saddle-Case Caddis
    American Grannom
      (Family: Brachycentridae, Genus: Brachycentrus)
      Tube-Case Caddis
    Little Western Weedy-Water Sedge
      (Family: Brachycentridae, Genus: Amiocentrus)
      Tube-Case Caddis
    Fall Caddis
      (Family: Limnephilidae, Genus: Dicosmoecus)
      Tube-Case Caddis
  Stoneflies, Midges
    Giant Salmonfly
      (Family: Pteronarcyidae, Genus: Pteronarcys)
    Golden Stoneflies
      (Family: Perlidae, Genus: Hesperoperla or Calineuria)
    Little Brown Stoneflies
      (Family: Nemouridae, Genus: Nemoura)
    Little Green Stoneflies
      (Family: Chloroperlidae, Genus: Alloperla or Sweltsa)
    Little Yellow Stoneflies
      (Family: Perlodidae, Genus: Isoperla)
    Midges
      (Family: Chironomidae)
  Fly Patterns

Hatch Guide For New England Streams by Thomas Ames, Jr.

 Order Stage Level Illustration Imitation      Other

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• • • •   • •     •     • • • • NE 271 • 2000
Topic organization:

  Introduction: The Madness Behind the Method
  The Life of a New England Freestone Stream
  Presentation on New England Streams
    Presenting a Wet Fly
    Presenting a Nymph
    Presenting a Dry Fly
    Tackling Tackle
    Presenting a Wet Fly
  A Short Primer on Aquatic Insect Structure
  Mayflies (Order Ephemeroptera)
    Tiny Fork-Gilled Nymphs (Genus Paraleptophlebia)
    Slate Winged Mahogany Dun (Genus Paraleptophlebia)
    Two Tailed Clinger - Nymph (Genus Epeorus)
    Quill Gordon - Dun (Epeorus pleuralis)
    Quill Gordon - Spinner (Epeorus pleuralis)
    Gray Winged Yellow Quill - Dun (Epeorus vitreus)
    Salmon Spinner (Epeorus vitreus)
    Crawlers - Nymphs (Genus Ephemerella)
    Hendrickson / Red Quill (Ephemerella subvaria)
    Red Quill Spinner (Ephemerella subvaria)
    Pale Evening Dun (Ephemerella invaria, rotunda)
    Pale Evening Spinner (Ephemerella invaria, rotunda)
    Sulphur - Dun (Ephemerella dorothea)
    Sulphur - Spinner (Ephemerella dorothea)
    Common Clingers, Nymphs
     (Genera Stenonema, Stenacron)
    American March Brown - Dun (Stenonema vicarium)
    American March Brown - Spinner
     (Stenonema vicarium)
    Cream Cahill - Dun
     (Stenonema mediopunctatum, modestum, others)
    Cream Cahill - Spinner
     (Stenonema mediopunctatum, modestum, others)
    Light Cahill - Dun (Genus Stenacron)
    Leadwing Coachman - Nymph (Genus Isonychia)
    Leadwing Coachman - Dun (Genus Isonychia)
    White Gloved Howdy - Spinner (Genus Isonychia)
    Gray Drake - Spinner (Genus Siphlonurus)
    Common Burrowers Nymphs (Family Ephemeridae)
    Green Drake - Dum (Ephemera guttulata)
    Coffin Fly - Spinner (Ephemera guttulata)
    Brown Drake - Spinner (Ephemera simulans)
    Yellow Drake - Spinner (Ephemera varia)
    Great Leadwinged Drake - Dun (Genus Hexagenia)
    Fringe Gilled Nymph (Anthopotamus distinctus)
    Golden Drake - Dun (Anthopotamus distinctus)
    Blue Winged Olives - Nymphs (Genus Drunella)
    Blue Winged Olives - Duns (Genus Drunella)
    Blue Winged Olives - Spinners (Genus Drunella)
    Dark Clingers - Nymphs (Genus Leucrocuta)
    Little Yellow Quill - Spinner (Genus Leucrocuta)
    White Fly - Nymph (Ephoron leukon, album)
    White Fly - Dun/Spinner (Ephoron leukon, album)
    Tiny Swimming Nymphs (Family Baetidae)
    Tiny Blue Winged Sulphurs, Rusts & Olives
     - Duns (Family Baetidae)
     - Spinners (Family Baetidae)
    Trico - Duns (Genus Tricorythodes)
    Trico - Spinners (Genus Tricorythodes)
  Caddisflies (Order Trichoptera)
    American Grannom
     - Larva (Genus Brachycentrus)
     - Pupa (Genus Brachycentrus)
     - Adult (Genus Brachycentrus)
    Saddle Case Maker
     - Pupa (Family Glossosomatidae)
     - Adult (Family Glossosomatidae)
    Common Net Spinners
     - Larvae (Family Hydropsychidae)
     - Pupae (Genera
       Hydropsyche, Ceratopsyche, Cheumatopsyche)
     - Adults (Genera
       Hydropsyche, Ceratopsyche, Cheumatopsyche)
    Cinnamon / Spotted Caddises
     - Adults (Genera Hydropsyche, Ceratopsyche)
    Little Sister Caddis - Adult (Genus Cheumatopsyche)
    Dark Blue Sedge - Larva (Psilotreta labida, frontalis)
    Dark Blue Sedge - Pupa (Psilotreta labida, frontalis)
    Dark Blue Sedge - Adult (Psilotreta labida, frontalis)
    Green Caddis - Larva (Family Rhyacophilidae)
    Green Caddis - Adult (Family Rhyacophilidae)
    "Alder Fly" - Larva (Macrostemum zebratum)
    "Alder Fly" - Pupa (Macrostemum zebratum)
    "Alder Fly" - Adult (Macrostemum zebratum)
    Northern Casemakers - Larvae (Family Limnephilidae)
    Northern Casemakers - Pupae (Family Limnephilidae)
    Northern Casemakers - Adults (Family Limnephilidae)
  Stoneflies (Order Plecoptera)
    Early Dark Stonefly Creepers
     (Family Taeniopterygidae)
    Early Dark Stonefly Adults (Family Taeniopterygidae)
    Little Green Stonefly Adults (Family Chloroperlidae)
    Giant Stoneflies - Creepers (Family Pteronarcyidae)
    Golden Stoneflies - Creepers (Family Perlidae)
    Golden Stoneflies - Adults (Family Perlidae)
    Little Yellow Stoneflies - Creepers (Family Perlodidae)
    Little Yellow Stoneflies - Adults (Family Perlodidae)
  Otheroptera: Damsels, Dragons, Others
    Dragonfly Larvae
     (Order Odonata; Suborder Anisoptera)
    Dragonfly Larvae
     (Order Odonata; Suborder Zygoptera)
    True Flies - Larvae (Order Diptera
    Midges - Adults (Order Diptera;
      Families Dixae, Chironomidae, others)
    Hellgrammites and Fishflies
     (Order Megaloptera; Family Corydalidae)
    Ants and Diving Wasps - Adults (Order Hymenoptera)
    Grasshoppers - Adults (Order Orthoptera)
  Fly Patterns
    Mayflies
    Caddisflies
    Stoneflies
    Other Orders

Hatches II by Al Caucci & Bob Nastasi

 Order Stage Level Illustration Imitation      Other

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•         • •     • • • • • • • NA 335 • 1986
Topic organization:

  An Introduction to The Mayfly
    The Mayfly Trout Relationship
    The Evolution of the Mayfly
    The Life Cycle of the Mayfly
    The Anatomy of the Mayfly
    Phonetics
    Glossary
  The Hatches
    Quick Reference Emergence Charts
     -Eastern and Midwestern North America
    Quick Reference Emergence Charts - West
    Family Concept of Identification
    The Basic Nymph Types
    The Family Tree
    The Crawlers (Family: Ephemerellidae, Caenidae,
     Tricorythidae, and Baetiscidae)
      Family Ephemerellidae (The Super Crawlers
        Genus Ephemerella
          E. subvaria
          E. invaria and rotunda
          E. dorothia
          E. infrequens and inermis
          E. cornuta
          E. lata
          E. glacialis
          E. flavilinea
          E. attenuata
      Family Leptophlebiidae (The Feeble-legged Crawlers
        Genus Paraleptophlebia
          P. adoptiva
        Genus Leptophlebia
          Leptophlebia Species
      Families Tricorythidae and Caenidae (The Tiny
       Crawlers)
        Genus Tricorythodes
          Tricorythodes Species
        Genus Caenis
        Genus Brachycercus
      Family Baetiscidae
    The Clingers
      Family Heptageniidae
        Genus Stenonema and Stenacron
          S. vicarium
          S. fuscum
          S. ithica and interpunctatum canadense
          Late Season Minor Species of Stenonema and
           Stenacron
        Genus Epeorus
          E. pleuralis
          Epeorus Species
        Genus Heptagenia
        Genus Rhithrogena
        Genus Cinygmula
    The Burrowers
      Families Ephemeridae, Potamanthidae and
       Polymitarcyidae (The Silt-dwelling Burrowers)
        Genus Ephemera
          E. guttulata
          E. varia
          E. simulans
        Genus Hexagenia and Litobrancha
          H. limbata
          Litobrancha recurvata
        Genus Potamanthus
          P. distinctus
        Genus Ephoron
    The Swimmers
      Family Baetidae, Siphlonuridae and Metretopodidae
       (The Darting Swimmers)
        Genus Baetis
          Baetis Species
        Genus Pseudocloeon and Cloeon
        Genus Callibaetis
        Genus Centroptilum
        Genus Isonychia
          I. bicolor
          I sadleri
        Genus Siphlonurus
        Genus Ameletus
        Genus Siphloplecton
      What Makes A Stream Tick
        Stream Research
      Imitation
        Fly Pattern Rationale
          The compara-nymphs and Deerhair Emerger
           Patterns
          The Indestructable Compara-dun
        Compara-Spinner and Spinner-falls
        Hook Rationale
        Compromise Fly Pattern Charts
        Specific Dressings

Mayflies by Ted Fauceglia

 Order Stage Level Illustration Imitation      Other

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•         • •     •     • •   • E/M 196 • 2005
Topic organization:

  MAYFLY METAMORPHOSIS
    Emerging Blue Quill Dun
    Molting Yellow Drake Dun
  THE MAYFLIES
    Blue-Winged Olives
      Baetis sp.
    Blue Quills
      Paraleptophlebia adoptiva
    Quill Gordon
      Epeorus pleuralis
    Hendrickson
      Ephemerella subvaria
    March Brown and Gray Fox
      Maccaffertium vicarium
    Small Eastern Blue-Winged Olive
      Acentrella attenuata
    Great Eastern Sulfur
      Ephemerella invaria
    Miniature Eastern Sulphur
      Ephemerella dorothea
    Miniature Blue-Winged Olive
      Acentrella sp.
    Gray Drake
      Siphlonurus sp.
    Eastern Green Drake
      Ephemera guttulata
    Brown Drake
      Ephemera simulans
    Yellow Drake
      Ephemera varia
    Great Olive-Winged Drake
      Hexagenia limbata
    Dark Green Drake
      Litobrancha recurvata
    Gray-Winged Yellow Quill
      Epeorus vitreus
    Eastern Blue-Winged Olive
      Drunella sp.
    Cream Cahills and Light Cahills
      Maccaffertium sp., Stenonema sp., Stenacron sp.
    Slate Drake
      Isonychia bicolor
    Golden Drake
      Anthopotamus sp.
    Tricos
      Tricorythodes sp.
    White Fly
      Ephoron sp.

Mayflies of Michigan Trout Streams by Justin W. Leonard and Fannie A. Leonard

 Order Stage Level Illustration Imitation      Other

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•         • •     • • • •     • MI 139 • 1962
Topic organization:

  Introduction
    Life Cycle
    Distribution
    Economic Importance
    Importance to Anglers
    Illustrations: Methods Followed
    Collecting, Preserving, and Rearing
    Classification
    Keys to Families and Genera
  MAYFLIES OF MICHIGAN TROUT STREAMS:
   DESCRIPTION OF SPECIES
    FAMILY EPHEMERIDAE
      Ephemera
        simulans
      Hexagenia
        atrocaudata, limbata, rigida, recurvata
    FAMILY CAENIDAE
      Genus Tricorythodes
        stygiatus
      Genus Brachycercus
        lacustris
      Genus Caenis
        simulans, jocosa, forcipata
    FAMILY EPHEMERELLIDAE
      Genus Ephemerella
        lata, walkeri, cornuta, deficiens, needhami,
        excrucians, dorothea, subvaria, invaria,
        rotunda, lutulenta, temporalis, simplex
    FAMILY BAETISCIDAE
      Genus Baetisca
        laurentina, obesa
    FAMILY LEPTOPHLEBIIDAE
      Genus Leptophlebia
        nebulosa, cupida
      Genus Paraleptophlebia
        debilis, praepedita, adoptiva, mollis, strigula
    FAMILY BAETIDAE
      Genus Siphlonurus
        rapidus, alternatus, quebecensis
      Genus Isonychia
        bicolor, sadleri, harperi
      Genus Callibaetis
        ferrugineus
      Genus Centroptilum
        album, convexum, rurostrigatum
      Genus Baetis,
        spinosus, pygmaeus, vagans, levitans, cingulatus,
        brunneicolor, hiemalis, intercalaris, phoebus
      Genus Pseudocloeon
        anoka
      Genus Cloeon
    FAMILY AMETROPIDAE
      Genus Metretopus
        norvegicus
      Genus Siphloplecton
        basale
    FAMILY HEPTAGENIIDAE
      Genus Stenonema
        canadense, frontale, interpunctatum, heterotarsale,
        tripunctatum, vicarium, fuscum, ithaca
  Glossary

Mayflies the Angler and the Trout by Fred L. Arbona, Jr.

 Order Stage Level Illustration Imitation      Other

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•         • •     • • • • • • • NA 188   1989
Topic organization:

  Mayfly Recognition and Imitation
  The Aquatic Fauna of a Trout Stream
  The Rationale of Mayfly Existence and Emergence
  The Mechanics of Good Presentation
  The Continuous Emergences of the Little Quills
   (Family Baetidae)
  Hatches of the Clingers: Gordon Quills, American March)
   Browns, Light Cahills, and Western Red Quills
   (Family Heptageniidae)
  The Early and Late-Season Hatches of Black Quills
   and Slate-Winged Mahogany Duns
   (Family Leptophlebiidae)
  Hendricksons, Pale Duns, and Blue Winged Olives
   (Family Ephemerellidae)
  Mid-Season Drakes of Placid Waters
   (Family Ephemeridae)
  All-Summer Hatches of Great Lead-Winged Coachmen
   and Gray Drakes
   (Family Siphlonuridae)
  The Late-Season Tiny White-Winged Tricos
   and Caenis Flies
   (Families Tricorythidae and Caenidae)
  Fly Tying Instructions
    Nymph Procedures
      Conventional Nymph
      Floating Nymph
      Flat-Body Nymph
      Streamlined Nymph
      Wiggle Nymph
    Dry Fly Instructions
      Emerging Dun
      The No-Hackle Fly
      Hen-Winged Dun
      Paradrake
      Poly or Hen-Winged Spinner
    Millimeter to Hook Type and Size Chart
  Advanced Entomology and Research
    Doing Your Own Research
    Equipment
    Nymph Identification
    Dry Form Identification
    Raising Mayflies
    Photography
    Nymph Identification
      Key to Species: Nymphs
    Dun (Subimago) and Spinner (Imago) Identification
      Key to Species: Duns
      Key to Species: Spinners

Mayflies, An Angler’s Study of Trout Water Ephemeroptera by Malcolm Knopp and Robert Cormier

 Order Stage Level Illustration Imitation      Other

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P
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P
h
o
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a
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s		 P
r
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t
a
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g
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.

D
a
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e		 G
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.

C
o
v
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a
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e		  #
 
 P
 a
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v
a
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e
 
N
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u
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l
i
s
h
e
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•         • •     • • • • • • • NA 336 • 1997
Topic organization:

  The Mayfly
    Biological Classification - The Systema Naturae
    Biological Classification of the Mayfly
    The Life Cycle of the Mayfly
    Mayfly Nymph Classification
    The Mayfly Nymph - An Anatomy
    The Mayfly Adult - An Anatomy
  The Mayfly and Its Imitation
    The beginnings of Modern Fly Fishing
    The Mayfly Nymph
    The Mayfly Emerger
    Cripples and Stillborns
    The Mayfly Dun - The Dry Fly
    The Mayfly Spinner
  The Swimming Nymphs
    The Swimming Nymphs - An Introduction
    The Blue-Winged Olives
     -Baetis, Diphetor, Labiobaetis and Acerpenna sp.
    The Tiny Blue-Winged Olives
     -Baetis and Acentrella sp.
    The Speckled Duns
     -Callibaetis sp.
    The Tiny Sulphur Duns
     - Centroptilum and Procloeon sp.
    The Gray-Winged Olive Quills
     - Centroptilum and Procloeon sp.
    The Gray Drakes
     - Siphlonurus sp.
    The Slate Drakes or Leadwings
     - Isonychia sp.
    The Brown Duns
     - Ameletus sp.
    The Great Speckled Olive Dun
     - Siphloplecton basale
  The Burrowing Nymphs - An Introduction
    The Brown Drake;
     - Ephemera simulans
    The Eastern Green Drake
     - Ephemera guttulata
    The Yellow Drake
     - Ephemera varia
    The Great Olive-winged Drake or Michigan Caddis
     - Hexagenia limbata and related sp.
    The Dark Green Drake
     - Litobrancha recurvata
    The White Drakes
     - Ephoron album and leukon
    The Golden Drakes
     - Anthopotamus sp.
  The Clinger Nymphs - An Introduction
    The Western March Browns
     - Rhithrogena sp.
    The Pale Evening Duns and Gray Fox
     - Heptagenia, Leucrocuta and Nixe sp.
    The Slate Duns and Quill Gordon
     - Epeorus sp.
    The Slate Maroon Drake
     - Ironodes nitidus
    The Dark Red Quills
     - Cinygmula sp.
    The Western Light Cahills
     - Cinygma sp.
    The American March Browns
     - Stenonema sp.
    The Light Cahills
     - Stenacron sp.
  The Crawler Nymphs - An Introduction
    The Eastern Hendrickson and Red Quill
     - Ephemerella subvaria
    The Eastern Sulphur Duns
     - Ephemerella invaria and rotunda
    The Eastern Pale Evening Dun
     - Ephemerella dorothea
    The Little Dark Hendrickson
     - Ephemerella needhami
    The Blue-Winged Olive
     - Ephemerella excrucians
    The Western Pale Morning Duns
     - Ephemerella infrequens, inermis, lacustris, mollita
    The Eastern Blue-Winged Olives
     - Drunella cornuta, walkeri, cornutella, longicornis
    The Western Green Drakes
     - Drunella grandis, doddsi, and subspecies
    The Small Western Green Drakes
     - Drunella flavilinea, coloradensis, and spinifera
    The Small Eastern Blue-Winged Olives
     - Attenella attenuata
    The Little Blue-Winged Olives
     - Attenella margarita
    The Small Blue-Winged Olives
     - Timpanoga simplex and lita
    The Great Red Quill
     - Timpanoga hecuba
    The Little Dark Hendricksons
     - Serratella deficiens and serrata
    The Small Western Dark Hendrickson
     - Serratella tibialis
    The Genus Caudatella
    The Chocolate Duns
     - Eurylophella sp.
    The Black Quills
     - Leptophlebia sp.
    The Blue Quills
     - Paraleptophlebia sp.
    The Tricos
     - Tricorythodes sp.
    The White-Winged Sulphurs
     - Caenis sp.
    The Genus Brachycercus
    The Genus Baetisca
  The Scientific and Common Naming of Mayflies
  Mayfly Hatch Charts
  A Fly Fisher's Aid to Mayfly Identification to the Genus
   Level
  A glossary of Entomological Terms
  The Phonetics of Latinized Mayfly sp. Names
  The Trout

Nymphs by Ernest Schwiebert

 Order Stage Level Illustration Imitation      Other

M
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I
n
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v
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r
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s		 A
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y		 G
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u
s		 S
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e
c
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s		 D
r
a
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i
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s		 B
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P
h
o
t
o
s		 C
o
l
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P
h
o
t
o
s		 P
a
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t
e
r
n
s		 P
r
e
s
e
n
t
a
t
i
o
n		 E
m
e
r
g
e
.

D
a
t
e		 G
e
o
.

C
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e
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a
g
e		  #
 
 P
 a
 g
 e
 s		 A
v
a
i
l
a
b
l
e
 
N
e
w		 P
u
b
l
i
s
h
e
d
• • • •   •       • •     • • • NA 339   1973
Topic organization:

  The Origins of Nymph Fishing
  The Evolution of Modern Nymph Tactics
  American Studies in Nymph Fishing
  Some Notes on Nymph-Fishing Tackle
  Fishing the Giant Stonefly Nymphs
  Notes on the Smaller Plecoptera
  Matching the Emerging Trichoptera
  Fishing Larval Imitations of the Caddis-Flies
  Some Notes on Fishing the Microcaddis
  Larvae and Pupae of the Larger Two-Winged Diptera
  Tiny Two-Winged Diptera and the Smutting Rise
  Fishing the Slow-Water Hemiptera
  Some Secrets of the Subaquatic Coleoptera
  Fishing The Freshwater Crustaceans
  The Larval Forms of the Sialidae
  The Fat-Bodied Nymphs of the Anisoptera
  The Slender-Bodied Nymphs of the Zygoptera
  The Gravel-Burrowing Ephemera Nymphs
  The Silt-Burrowing Hexagenia Nymphs
  The Fringe-Gilled nymphs of the Potamanthus
  The Swimming Nymphs of the Siphlonurus and
   Siphloplecton
  The Tiny Swimming Nymphs of the Baetis Flies
  The Clambering Slow-Water Callibaetis Nymphs
  The Fast-Water Swimming Isonychias
  The Thick-Gilled Leptophlebia Nymphs
  The Tiny Fork-Gilled Paraleptophlebia Nymphs
  Minute Nymphal Forms of the Caenidae
  The Minute Cloeon, Neocloeon, and the Pseudocloeon
   Flies
  The Tiny Swimming Nymphs of the Centroptilum Hatches
  Fishing the Two-Tailed Epeorus Nymphs
  The Clinging Plate-Gilled Rhithrogenas
  Clinging Nymphs of the Minor Heptageniidae
  The Nymphs of the Swift-Water Stenonema Flies
  The Humpbacked Nymphs of the Baetisca Flies
  The Ubiquitous Nymphs of the Ephemerella Flies
  

Selective Trout by Doug Swisher and Carl Richards

 Order Stage Level Illustration Imitation      Other

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r
a
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s		 B
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P
h
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t
o
s		 C
o
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P
h
o
t
o
s		 P
a
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t
e
r
n
s		 P
r
e
s
e
n
t
a
t
i
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n		 E
m
e
r
g
e
.

D
a
t
e		 G
e
o
.

C
o
v
e
r
a
g
e		  #
 
 P
 a
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a
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N
e
w		 P
u
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h
e
d
• • •     • •     • • • • •   • NA 184 • 1971
Topic organization:

  Science and Selectivity
  Collecting, Photographing, Identifying
  The Need for Realistic Imitation
  Mayflies - The Upwings
  Patterns Evolved - No Hackle Duns, Paraduns,
   Emerging Patterns, Spinners
  Streamside Procedure
  The early Season
  The Midseason
  The Late Season
  Caddisflies, Stone Flies, Midges - The Downwings
  Terrestrials
  The "Super Hatches"
  Night Fishing
  A Barbless Hook
  Identification Of Mayflies
    Structure of Generalized Mayfly Nymphs
    Structure of Generalized Mayfly Adult Male
    Species Descriptions
  Millimeter to Hook Size Guide

The Caddisfly Handbook by Dick Pobst and Carl Richards

 Order Stage Level Illustration Imitation      Other

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P
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P
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r
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s
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t
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.

D
a
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e		 G
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C
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 P
 a
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N
e
w		 P
u
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l
i
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h
e
d
  •       • •     • •   • •   • NA 133 • 1998
Topic organization:

  Introduction
    Finding the Really Important Hatches
    The Importance of Behavior
    Scientific Names or Common Names
  The Caddis Rivers
    Tailwaters
    Trout Streams
    Spring Creeks
    Not Two Rivers Are Alike
    The Muskegon
  The Life Cycle of the Caddisfly
  Identifying the Caddisflies
    First, Identify the Larvae or Cases
    Second, Identify the Adults
  Major Caddisflies West of the Mississippi
    Mother's Day Caddis - Brachycentrus
    Spotted Sedge - Ceratopsyche/Hydropsyche
    Little Sister Sedge - Cheumatopsyche
    Short-Horn Sedge - Glossosoma
    Weedy Water Sedge - Micrasema/Amiocentrus
    Speckled Peter - Helicopsyche
    Green Sedge - Rhyacophila
    Little Brown Sedge - Lepidostoma
    Gray-Winged Long-Horn Sedge - Oecetis
    Black Dancer - Mystacides
    Great Gray-Spotted Sedge - Arctopsyche
    Great Silver Striped Sedge - Hesperophylax
    Giant Orange Sedge - Dicosmoecus
    Trout Lake Caddis
  Major Hatches East of the Mississippi
    Black Caddis - Brachycentrus
    Cinnamon Caddis - Ceratopsyche
    Little Olive Caddis - Cheumatopsyche
    Dark Blue Sedge - Psilotreta
    Green Sedge - Rhyacophila
    Short-Horn Sedge - Glossosoma
    Tiny Black Caddis - Micrasema
    Zebra Caddis - Macrostemum
    Tan-Winged Long-Horn Sedge - Oecetis
    Dark Long-Horn Sedge - Ceraclea
    White Miller - Nectopsyche
    Dot-Winged Sedge - Frenesia
    Great Brown Autumn Sedge - Pycnopsyche
  Tying Caddis Imitations
  Master List of Caddis Patterns
  Occasionally Important Caddisflies
  Fisherman's Simplifies Keys

The Complete Book Of Western Hatches: An Anglers Entomology and Fly Pattern Field Guide by Rick Hafele and Dave Hughes

 Order Stage Level Illustration Imitation      Other

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a
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r
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s		 B
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P
h
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P
h
o
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o
s		 P
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n
s		 P
r
e
s
e
n
t
a
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i
o
n		 E
m
e
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g
e
.

D
a
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e		 G
e
o
.

C
o
v
e
r
a
g
e		  #
 
 P
 a
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 e
 s		 A
v
a
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a
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l
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N
e
w		 P
u
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l
i
s
h
e
d
• • • •   • •   •   • • • • • • W 223 • 1981
Topic organization:

  KNOWING THE ORDERS OF INSECTS
  MAYFLIES
   (ORDER: EPHEMEROPTERA}
  STONEFLIES
   (ORDER: PLECOPTERA)
  DRAGONFLIES AND DAMSELFLIES
   (ORDER: ODONATA)
  WATERBOATMEN AND BACKSWIMMERS
   (ORDER: HEMIPTERA)
  CADDISFLIES
   (ORDER: TRICHOPTERA)
  ALDERFLIES AND DOBSONFLIES
   (ORDER: MEGALOPTERA)
  WATER BEETLES
   (ORDER: COLEOPTERA)
  TRUE FLIES
   (ORDER: DIPTERA)
  GLOSSARY

The Hatches Made Simple by Charles R. Meck

 Order Stage Level Illustration Imitation      Other

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P
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P
h
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s		 P
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t
a
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m
e
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g
e
.

D
a
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e		 G
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o
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C
o
v
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a
g
e		  #
 
 P
 a
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a
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N
e
w		 P
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h
e
d
• • •     • •     • • • • • • • NA 261 • 2002
Topic organization:

  What Trout Eat
  Spinners
  Downwings
  An Introduction to the Hatches
  Early Spring - Grays
  Summer Hatches in the Morning and Afternoon - Olives
   and a Few Dark Grays and Browns
  Summer Evenings - Those Cream Mayflies
  Summer Evenings - Those Dark Hatches
  Early Fall and Winter - The Grays (and Olives) Return
  Patterns for the Hatches
  Special Considerations
  A Final Hatch
  Mayfly Identification
  Glossary

Western Mayfly Hatches: From the Rockies To The Pacific by Rick Hafele and Dave Hughes

 Order Stage Level Illustration Imitation      Other

M
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I
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I
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P
h
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P
h
o
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s		 P
r
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s
e
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t
a
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i
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m
e
r
g
e
.

D
a
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e		 G
e
o
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C
o
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a
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 P
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N
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•         • •   •   •   • • • • W 267 • 2004
Topic organization:

  Matching Mayflies
    A Philosophy for Matching Hatches
    Collecting and Observing Mayflies
    Recognizing Mayflies
    About Mayflies
    Timing and Importance
  Mayfly Swimmers
    Baetis (Blue-Winged Olives)
    Callibaetis (Speckle-Wings)
    Siphlonurus (Gray Drakes)
    Ameletus (Brown Duns)
    Isonychia (Great Western Leadwings)
  Mayfly Crawlers
    Ephemerella (Pale Morning Duns)
    Serratella (Small Western Dark Hendricksons)
    Smaller Drunella (Flavs, Lesser Green Drakes)
    Greater Drunella (Western Green Drakes)
    Attenella (Small Blue-Winged Olives)
    Caudatella (No Common Name)
    Timpanoga (Great Blue-Winged Red Quills)
    Paraleptophlebia (Mahogany Duns)
    Leptophlebia (Western Black Quills)
    Tricorythodes (Tricos)
    Caenis (White-Winged Sulphurs)
  Mayfly Clingers
    Rhithrogena (Western March Browns)
    Epeorus (Pink Lady, Slate Brown Duns)
    Heptagenia, Nixe, Leucrocuta, Cinygma
     (Pale Evening Duns)
    Cinygmula (Dark Red Quills)
  Mayfly Burrowers
    Hexagenia (Hex)
    Ephemera (Brown Drakes)
    Ephoron (White Drakes)
  Much About Mayflies
  Glossary

Which book to get?

They are all excellent.  Each has its own "treasure trove" of information.   However, as stated earlier, it is my hope that this article will have provided sufficient information for you to choose the book or books that best satisfy your particular needs.  However, if you are left with any questions, I would be more than happy to answer them.

Roger Rohrbeck, a flyfisher, fly tyer, and entomology enthusiast from the state of Washington can be contacted through his website, FlyfishingEntomology.com or on FlyTyingForum.com - Entomology Forum, where he participates as Taxon.

Created: 03/20/2016   Last modified: 03/20/2016    www.FlyfishingEntomology.com